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A very small woodland bird with a long tail, usually seen flitting about in the treetops, giving a short whining callnote. Often it darts out in a short, quick flight to snap up a tiny insect in mid-air. Widespread in summer, its breeding range is still expanding toward the north.

Until the late 1980s, this bird was regarded as just a local form of the Black-tailed Gnatcatcher. With its recognition as a full species, it also became an endangered species: its limited habitat along the southern California coast is being taken over by housing tracts and other developments. California Gnatcatchers live in coastal sage scrub, a low shrubby habitat that is also home to other specialized animals and plants.

This long-tailed little insect-eater is at home in the desert southwest, even in arid scrub and creosote bush flats where there are few other birds. Black-tailed Gnatcatchers live in pairs all year, foraging together actively in the low brush. They stay in contact with each other using a wide variety of calls; some of these calls sound suspiciously like imitations of other desert birds, such as Verdin or Black-throated Sparrow.

This small songbird from western Mexico has been flirting with the Arizona border since the early 1970s. It has appeared in many different canyons in southeastern Arizona, and it has been known to nest there a number of times. Its occurrence is still erratic, however, and it does not seem to become permanently established in any one locality. In feeding and nesting behavior, it is quite similar to our other gnatcatchers.